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- The hexagonal cells fit closely
together, and each bear two flagella.
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- The cells are connected by
cytoplasmic extensions.
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- Volvox is a motile green algal colony composed
of 500-50,000 cells arranged along the periphery of a hollow
mucilaginous sphere. These large colonies are often visible with
the naked eye. Each cell has its own individual mucilage that
may be distinct or inconspicuous. Strands of protoplasm hold
the cells together and are more readily seen in young colonies
of certain species. The colonial mucilage is surrounded by a
boundary layer that has a similar chemical composition to the
cell wall of Chlamydomonas.
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- Like Chlamydomonas,
each spherical or ovoid cell has two equal flagella with a pair
of contractile vacuoles at their base and a large cup-shaped
chloroplast with a single pyrenoid. The colonies coordinate the
flagellar movements of cells on different sides of the sphere
in order to swim with a rolling motion and change direction as
necessary. Volvox is phototactic and able to swim towards
moderate light.
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- Volvox colonies demonstrate polar organization.
The anterior cells have larger eyespots and greater phototactic
abilities, while the posterior cells are involved with reproduction.
Volvox is widely used by researchers as a genetic and
developmental model to further understand cellular differentiation.
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- The 20 Volvox species
are distinguished mainly by the characteristics of the mucilage
and the cytoplasmic strands connecting adjacent cells. DNA sequencing
has shown that the genus is polyphyletic and that the various
species have evolved from four ancestral lineages. Researchers
may eventually reclassify species into different genera to reflect
this relationship.
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